26:21a From the descendants of Ladan, who were Gershonites through Ladan

They were descendants of Ladan through ⌊Levi’s eldest son⌋ Gershon.
-or-
They belonged to the family of Ladan. Ladan was from the clan of Gershon.

26:21b and heads of the families of Ladan the Gershonite, were Jehieli,

The leaders of families in the clan of Ladan were Jehieli
-or-
Jehieli was the chief/leader of one of Ladan’s family groups.

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